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- Title
TAXONOMIC REVISION OF DYAKIA JANUS FROM PENINSULAR MALAYSIA (PULMONATA: DYAKIIDAE), WITH NOTES ON OTHER SINISTRALLY COILED HELICARIONOIDS.
- Authors
Sutcharit, Chirasak; Tongkerd, Piyoros; Aileen Tan, Shau-Hwai; Panha, Somsak
- Abstract
The taxonomic status of Dyakia janus (Beck, 1837), a poorly known Malayan land snail, is re-assessed and a neotype designated. Dyakia janus possesses a long epiphallus, but a fl agellum and penial verge are both absent. The species is also characterised by having a long gametolytic duct, an amatorial organ comprising three major glands and a calcifi ed, spiked papilla, a vagina surrounded by sponge-like tissue, a mantle collar that has a posterior right dorsal lobe, and a radula that exhibits unicuspid teeth. Two sinistral species previously misidentifi ed as true Dyakia, namely Helix retrorsa Gould, 1844 and Nanina salangana Martens, 1883, were compared with D. janus. We show that the former two species differ from true Dyakia s.s. in the absence of an amatorial organ gland and presence of a penial sheath, epiphallic caecum, fl agellum, and a gametolytic sac that is connected by a short duct to the vagina. The reproductive system is a signifi cant source of taxonomic characters for generic classifi cation, and these two species should rightly be assigned to the family Ariophantidae instead of Dyakiidae.
- Subjects
MALAYA; HELICARIONIDAE; SNAILS; ANIMAL classification; PULMONATA; ANIMAL species; COMPARATIVE studies
- Publication
Raffles Bulletin of Zoology, 2012, Vol 60, Issue 2, p279
- ISSN
0217-2445
- Publication type
Article